Oxygen-18 Enrichment of Planktonic Foraminifera Due to Gametogenic Calcification Below the Euphotic Zone

Abstract
Empty shells of spinose planktonic foraminifera on the seabed are significantly enriched in oxygen-18 as compared with the shells of their living counterparts in surface waters. This enrichment is due to gametogenic calcification, which extracts calcium carbonate from deeper and colder waters as the shell sinks below the euphotic zone. JEAN-CLAUDE DUPLESSY