Abstract
A series of papers, Hamilton & May (1977), Motro, 1982, Motro, 1983 and Frank (1986) have developed and generalized a model of dispersal of offspring to random sites in a stable environment. I present an inclusive fitness model for such dispersal behaviour, applicable in diploid or haplodiploid populations with dispersal under offspring or maternal control. This model provides an illustration of the general discussion of inclusive fitness models found in Taylor, 1988a, Taylor, 1988b. In particular, it provides a good example of the way in which relatedness coefficients can depend, not only on the genetic structure of the population, but on the mechanism of control of the behaviour under study.

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