Jurassic ammonites from Jebel Tuwaiq, central Arabia with stratigraphical introduction

Abstract

The Jurassic system as developed along a strike of some 500 miles in the Jebel Tuwaiq, central Saudi Arabia (Nejd), is described and divided into formations, of which the names are here published for the first time, with a synopsis of their lithology and fossil contents. Apart from small collections,'including only one ammonite species, made on camel journeys by H. St J. B. Philby, nothing was hitherto known of these formations; their age (except for the Middle Callovian date given by the ammonite), and the existence of most of them, were unknown until the area began to be mapped and explored geologically about 1940 by geologists of the Arabian American Oil Company, whose results are here summarized. The total thickness of marine Jurassic rocks described exceeds 1000 m., all of it in neritic facies. This sequence is divided, in ascending order, into Marrat, Dhruma, Tuwaiq Mountain, Hanifa, Jubaila and Riyadh formations, above which follows the Cretaceous system. Below the marine Marrat is the Minjur Sandstone (315 m.), of 'continental’ facies, with only obscure plant remains, which possibly corresponds to the supposedly Rhaetic-Lower Lias plant-bearing Kohlan Sandstones of the Yemen. The underlying Jilh formation of Saudi Arabia contains marine Middle Triassic fossils near the top. Collections of ammonites establish the presence of marine Jurassic stages from Lower Toarcian (Lower Marrat) to Lower Kimeridgian (Jubaila); the Riyadh has not yet yielded ammonites at outcrop. The ammonites described are of extraordinary interest from the points of view of palaeogeography, correlation, phylogeny and systematics. While most of the faunas contain just enough links with other parts of the world to establish broad correlations, many are entirely new, or confined, outside Arabia, to Sinai. In the Toarcian there was faunal continuity with Madagascar and Baluchistan, in the Bajocian only with Sinai, from the Middle Bathonian onwards with western Europe. Most of the commonest cosmopolitan genera are missing from central Arabia, where there lived a succession of highly peculiar forms characterized at several successive horizons by unstable suture-lines. It has been found necessary to make eight new genera, besides three already named but hitherto not found outside Sinai, Madagascar and Baluchistan, although employing a taxonomic scale so large that some palaeontologists will consider it old-fashioned. This is believed, however, to be more useful at this stage of our knowledge than a proliferation of generic names based on inadequate material; for the area from which the collections were made is about equal to that of all the Jurassic outcrops of England. The paper concludes with a discussion of problems of evolution, speciation, correlation, and palaeogeography raised by the material. In an appendix are described relevant Bajocian ammonites from Sinai, with a hitherto unpublished section of the strata.