Paleoanthropology Fossil hominins from South Africa are enriching the story of early human evolution and dispersal. Herries et al. describe the geological context and dating of the hominin-bearing infilled cave, or palaeocave, at a site called Drimolen in South Africa (see the Perspective by Antón). They focus on the age and context of a recently discovered Homo erectus sensu lato fossil and a Paranthropus robustus fossil, which they dated to ∼2.04 million to 1.95 million years ago. This makes Drimolen one of the best-dated sites in South Africa and establishes these fossils as the oldest definitive specimens of their respective species ever discovered. The age confirms that species of Australopithecus, Paranthropus , and early Homo overlapped in the karst of South Africa ∼2 million years ago. Science , this issue p. [eaaw7293][1]; see also p. [34][2] [1]: /lookup/doi/10.1126/science.aaw7293 [2]: /lookup/doi/10.1126/science.abb4590